diff options
Diffstat (limited to 'man/occuCOP.Rd')
-rw-r--r-- | man/occuCOP.Rd | 249 |
1 files changed, 249 insertions, 0 deletions
diff --git a/man/occuCOP.Rd b/man/occuCOP.Rd new file mode 100644 index 0000000..f0b5be0 --- /dev/null +++ b/man/occuCOP.Rd @@ -0,0 +1,249 @@ +\name{occuCOP} + +\alias{occuCOP} + +\encoding{UTF-8} + +\title{Fit the occupancy model using count dta} + +\usage{ +occuCOP(data, + psiformula = ~1, lambdaformula = ~1, + psistarts, lambdastarts, starts, + method = "BFGS", se = TRUE, + engine = c("C", "R"), na.rm = TRUE, + return.negloglik = NULL, L1 = FALSE, ...)} + +\arguments{ + + \item{data}{An \code{\link{unmarkedFrameOccuCOP}} object created with the \code{\link{unmarkedFrameOccuCOP}} function.} + + \item{psiformula}{Formula describing the occupancy covariates.} + + \item{lambdaformula}{Formula describing the detection covariates.} + + \item{psistarts}{Vector of starting values for likelihood maximisation with \code{\link{optim}} for occupancy probability \eqn{\psi}{psi}. These values must be logit-transformed (with \code{\link{qlogis}}) (see details). By default, optimisation will start at 0, corresponding to an occupancy probability of 0.5 (\code{plogis(0)} is 0.5).} + + \item{lambdastarts}{Vector of starting values for likelihood maximisation with \code{\link{optim}} for detection rate \eqn{\lambda}{lambda}. These values must be log-transformed (with \code{\link{log}}) (see details). By default, optimisation will start at 0, corresponding to detection rate of 1 (\code{exp(0)} is 1).} + + \item{starts}{Vector of starting values for likelihood maximisation with \code{\link{optim}}. If \code{psistarts} and \code{lambdastarts} are provided, \code{starts = c(psistarts, lambdastarts)}.} + + \item{method}{Optimisation method used by \code{\link{optim}}.} + + \item{se}{Logical specifying whether to compute (\code{se=TRUE}) standard errors or not (\code{se=FALSE}).} + + \item{engine}{Code to use for optimisation. Either \code{"C"} for fast C++ code, or \code{"R"} for native R code.} + + \item{na.rm}{Logical specifying whether to fit the model (\code{na.rm=TRUE}) or not (\code{na.rm=FALSE}) if there are NAs in the \code{\link{unmarkedFrameOccuCOP}} object.} + + \item{return.negloglik}{A list of vectors of parameters (\code{c(psiparams, lambdaparams)}). If specified, the function will not maximise likelihood but return the negative log-likelihood for the those parameters in the \code{nll} column of a dataframe. See an example below.} + + \item{L1}{Logical specifying whether the length of observations (\code{L}) are purposefully set to 1 (\code{L1=TRUE}) or not (\code{L1=FALSE}).} + + \item{\dots}{Additional arguments to pass to \code{\link{optim}}, such as lower and upper bounds or a list of control parameters.} + } + +\description{This function fits a single season occupancy model using count data.} + +\details{ + + See \code{\link{unmarkedFrameOccuCOP}} for a description of how to supply data to the \code{data} argument. See \code{\link{unmarkedFrame}} for a more general documentation of \code{unmarkedFrame} objects for the different models implemented in \pkg{unmarked}. + + \subsection{The COP occupancy model}{ + + \code{occuCOP} fits a single season occupancy model using count data, as described in Pautrel et al. (2023). + + The \strong{occupancy sub-model} is: + + \deqn{z_i \sim \text{Bernoulli}(\psi_i)}{z_i ~ Bernoulli(psi_i)} + + \itemize{ + \item With \eqn{z_i}{z_i} the occupany state of site \eqn{i}{i}. \eqn{z_i=1}{z_i = 1} if site \eqn{i}{i} is occupied by the species, \emph{i.e.} if the species is present in site \eqn{i}{i}. \eqn{z_i=0}{z_i = 0} if site \eqn{i}{i} is not occupied. + \item With \eqn{\psi_i}{psi_i} the occupancy probability of site \eqn{i}{i}. + } + + The \strong{observation sub-model} is: + + \deqn{ + N_{ij} | z_i = 1 \sim \text{Poisson}(\lambda_{ij} L_{ij}) \\ + N_{ij} | z_i = 0 \sim 0 + }{ + N_ij | z_i = 1 ~ Poisson(lambda_is*L_is) + N_ij | z_i = 0 ~ 0 + } + + \itemize{ + \item With \eqn{N_{ij}}{N_ij} the count of detection events in site \eqn{i}{i} during observation \eqn{j}{j}. + \item With \eqn{\lambda_{ij}}{lambda_ij} the detection rate in site \eqn{i}{i} during observation \eqn{j}{j} (\emph{for example, 1 detection per day.}). + \item With \eqn{L_{ij}}{L_ij} the length of observation \eqn{j}{j} in site \eqn{i}{i} (\emph{for example, 7 days.}). + } + + What we call "observation" (\eqn{j}{j}) here can be a sampling occasion, a transect, a discretised session. Consequently, the unit of \eqn{\lambda_{ij}}{lambda_ij} and \eqn{L_{ij}}{L_ij} can be either a time-unit (day, hour, ...) or a space-unit (kilometer, meter, ...). + } + + \subsection{The transformation of parameters \eqn{\psi} and \eqn{\lambda}}{ + In order to perform unconstrained optimisation, parameters are transformed. + + The occupancy probability (\eqn{\psi}) is transformed with the logit function (\code{psi_transformed = qlogis(psi)}). It can be back-transformed with the "inverse logit" function (\code{psi = plogis(psi_transformed)}). + + The detection rate (\eqn{\lambda}) is transformed with the log function (\code{lambda_transformed = log(lambda)}). It can be back-transformed with the exponential function (\code{lambda = exp(lambda_transformed)}). + } + +} + +\value{\code{unmarkedFitOccuCOP} object describing the model fit. See the \code{\linkS4class{unmarkedFit}} classes.} + +\references{ + +Pautrel, L., Moulherat, S., Gimenez, O. & Etienne, M.-P. Submitted. \emph{Analysing biodiversity observation data collected in continuous time: Should we use discrete or continuous-time occupancy models?} Preprint at \href{https://doi.org/10.1101/2023.11.17.567350}{https://doi.org/10.1101/2023.11.17.567350}. + +} + +\author{Léa Pautrel} + +\seealso{ + \code{\link{unmarked}}, + \code{\link{unmarkedFrameOccuCOP}}, + \code{\link{unmarkedFit-class}} +} + + +\examples{ +set.seed(123) +options(max.print = 50) + +# We simulate data in 100 sites with 3 observations of 7 days per site. +nSites <- 100 +nObs <- 3 + +# For an occupancy covariate, we associate each site to a land-use category. +landuse <- sample(factor(c("Forest", "Grassland", "City"), ordered = TRUE), + size = nSites, replace = TRUE) +simul_psi <- ifelse(landuse == "Forest", 0.8, + ifelse(landuse == "Grassland", 0.4, 0.1)) +z <- rbinom(n = nSites, size = 1, prob = simul_psi) + +# For a detection covariate, we create a fake wind variable. +wind <- matrix(rexp(n = nSites * nObs), nrow = nSites, ncol = nObs) +simul_lambda <- wind / 5 +L = matrix(7, nrow = nSites, ncol = nObs) + +# We now simulate count detection data +y <- matrix(rpois(n = nSites * nObs, lambda = simul_lambda * L), + nrow = nSites, ncol = nObs) * z + +# We create our unmarkedFrameOccuCOP object +umf <- unmarkedFrameOccuCOP( + y = y, + L = L, + siteCovs = data.frame("landuse" = landuse), + obsCovs = list("wind" = wind) +) +print(umf) + +# We fit our model without covariates +fitNull <- occuCOP(data = umf) +print(fitNull) + +# We fit our model with covariates +fitCov <- occuCOP(data = umf, psiformula = ~ landuse, lambdaformula = ~ wind) +print(fitCov) + +# We back-transform the parameter's estimates +## Back-transformed occupancy probability with no covariates +backTransform(fitNull, "psi") + +## Back-transformed occupancy probability depending on habitat use +predict(fitCov, + "psi", + newdata = data.frame("landuse" = c("Forest", "Grassland", "City")), + appendData = TRUE) + +## Back-transformed detection rate with no covariates +backTransform(fitNull, "lambda") + +## Back-transformed detection rate depending on wind +predict(fitCov, + "lambda", + appendData = TRUE) + +## This is not easily readable. We can show the results in a clearer way, by: +## - adding the site and observation +## - printing only the wind covariate used to get the predicted lambda +cbind( + data.frame( + "site" = rep(1:nSites, each = nObs), + "observation" = rep(1:nObs, times = nSites), + "wind" = getData(fitCov)@obsCovs + ), + predict(fitCov, "lambda", appendData = FALSE) +) + +# We can choose the initial parameters when fitting our model. +# For psi, intituively, the initial value can be the proportion of sites +# in which we have observations. +(psi_init <- mean(rowSums(y) > 0)) + +# For lambda, the initial value can be the mean count of detection events +# in sites in which there was at least one observation. +(lambda_init <- mean(y[rowSums(y) > 0, ])) + +# We have to transform them. +occuCOP( + data = umf, + psiformula = ~ 1, + lambdaformula = ~ 1, + psistarts = qlogis(psi_init), + lambdastarts = log(lambda_init) +) + +# If we have covariates, we need to have the right length for the start vectors. +# psi ~ landuse --> 3 param to estimate: Intercept, landuseForest, landuseGrassland +# lambda ~ wind --> 2 param to estimate: Intercept, wind +occuCOP( + data = umf, + psiformula = ~ landuse, + lambdaformula = ~ wind, + psistarts = rep(qlogis(psi_init), 3), + lambdastarts = rep(log(lambda_init), 2) +) + +# And with covariates, we could have chosen better initial values, such as the +# proportion of sites in which we have observations per land-use category. +(psi_init_covs <- c( + "City" = mean(rowSums(y[landuse == "City", ]) > 0), + "Forest" = mean(rowSums(y[landuse == "Forest", ]) > 0), + "Grassland" = mean(rowSums(y[landuse == "Grassland", ]) > 0) +)) +occuCOP( + data = umf, + psiformula = ~ landuse, + lambdaformula = ~ wind, + psistarts = qlogis(psi_init_covs)) + +# We can fit our model with a different optimisation algorithm. +occuCOP(data = umf, method = "Nelder-Mead") + +# We can run our model with a C++ or with a R likelihood function. +## They give the same result. +occuCOP(data = umf, engine = "C", psistarts = 0, lambdastarts = 0) +occuCOP(data = umf, engine = "R", psistarts = 0, lambdastarts = 0) + +## The C++ (the default) is faster. +system.time(occuCOP(data = umf, engine = "C", psistarts = 0, lambdastarts = 0)) +system.time(occuCOP(data = umf, engine = "R", psistarts = 0, lambdastarts = 0)) + +## However, if you want to understand how the likelihood is calculated, +## you can easily access the R likelihood function. +print(occuCOP(data = umf, engine = "R", psistarts = 0, lambdastarts = 0)@nllFun) + +# Finally, if you do not want to fit your model but only get the likelihood, +# you can get the negative log-likelihood for a given set of parameters. +occuCOP(data = umf, return.negloglik = list( + c("psi" = qlogis(0.25), "lambda" = log(2)), + c("psi" = qlogis(0.5), "lambda" = log(1)), + c("psi" = qlogis(0.75), "lambda" = log(0.5)) +)) +} + +\keyword{models} |